Knuckles debuted in 1994’s Sonic the Hedgehog 3 for the Sega Genesis. Created by designer Takashi Yuda, he was designed as a physical opposite of Sonic, a strong, grounded character to serve as ...

Throbbing, aching, or shooting pain in the knuckles can result from an injury or an underlying medical condition, such as arthritis or a cyst. The treatment for knuckle pain may depend on the ...