Those two question marks at linebacker became defensive exclamation points by the end of the season, combining for 279 sacks, including 20 for losses, 6 1/2 sacks and an interception. “You had Oren Burks who we had just signed,” Fangio said.

Landon Dickerson (knee) and Ben VanSumeren (ACL), who were absent during last week’s viewing of OTAs, both worked out on Tuesday. Lane Johnson and Reed Blankenship also returned. There were four absences: DeVonta Smith, who also missed last week’s session, Johnny Wilson, Elijah Cooks and E.J. Jenkins.

The Philadelphia Eagles held their first OTAs practice open to the media on Wednesday. Here are five takeaways.